Mark Harvey

Qualifications

B.Sc., Ph.D.

Research

Systematics of arachnids and myriapods, including their phylogenetic history and evolutionary relationships.
Taxonomy of arachnids and myriapods, and especially the documentation and description of new species. To date, he has described nearly 900 new species and more than 90 new genera.

Collections

Researches the taxonomy and systematics of arachnids and myriapods through the collections.
Develops the Museum’s collections through targeted fieldwork and collaborations with other organisations, including government, universities and industry.

Services (to other bodies)

Represents the Museum on committees and panels providing advice to government.

Exhibitions

Provides identifications and advice for environmental consultants, industry and government agencies.
Regularly presents talks and provides media coverage of work on arachnids and myriapods.
Editor of the Australian Journal of Taxonomy; Associate Editor of Invertebrate Systematics. Member of the editorial board of several other scholarly journals.
Co-supervisor to several post-graduate students at local and international universities.
Member of the International Commission on Zoological Nomenclature: http://iczn.org/content/dr-mark-s-harvey

Websites

World Pseudoscorpiones Catalog: https://wac.nmbe.ch/order/pseudoscorpiones/3

Selected Publications

Mark’s full publication list can be found here

2024

Harms, D., Harvey, M. S., Roberts, J. D. & Loria, S. F. (2024) Tectonically-driven climate change and the spread of temperate biomes: Insights from dragon pseudoscorpions (Pseudotyrannochthoniidae), a globally distributed arachnid lineage. Journal of Biogeography, 51, 1032–1048.

Harvey, M. S. & Steinmann, D. B. (2024) A new troglomorphic species of Larca (Pseudoscorpiones, Larcidae) from Colorado. ZooKeys, 1198, 279–294.

Hlebec, D., Harms, D., Kučinić, M. & Harvey, M. S. (2024) Integrative taxonomy of the pseudoscorpion family Chernetidae (Pseudoscorpiones: Cheliferoidea): evidence for new range-restricted species in the Dinaric Karst. Zoological Journal of the Linnean Society, 200, 644–669.

Rix, M. G. & Harvey, M. S. (2024) A new species of pelican spider (Araneae: Archaeidae) from the Whitsunday hinterland of central-eastern Queensland. Australian Journal of Taxonomy, 64, 1–7.

Sagastume-Espinoza, K. O., Simmons, L. W. & Harvey, M. S. (2024) Use of geometric morphometrics to distinguish trapdoor spider morphotypes (Mygalomorphae: Anamidae: Proshermacha): a useful tool for mygalomorph taxonomy. Journal of Arachnology, 52, 31–40.

Sherwood, D., Grignet, V., Harvey, M. S., Sharp, A., Wilkins, V., Ashmole, M. & Ashmole, P. (2024) David and Goliath: on the pseudoscorpions of Ascension Island, including the world’s largest, Garypus titanius Beier, 1961, and a new, minute, Neocheiridium Beier, 1932 (Arachnida: Pseudoscorpiones). Natura Somogyiensis, 42, 131–150.

Sloan, N. S., Harvey, M. S., Huey, J. A. & Simmons, L. W. (2024) Rapid divergent evolution of internal female genitalia and the coevolution of male genital morphology revealed by micro-computed tomography. Proceedings of the Royal Society B: Biological Sciences, 291, 20232883.

2023

Buzatto, B. A., Clark, H. L., Harvey, M. S. & Volschenk, E. S. (2023) Two new species of burrowing scorpions (Urodacidae: Urodacus) from the Pilbara region of Western Australia with identical external morphology. Australian Journal of Zoology, 71, 1–18.

Ceríaco, L. M. P., Aescht, E., Ahyong, S. T., Ballerio, A., Bouchard, P., Bourgoin, T., Dmitriev, D., Evenhuis, N., Grygier, M. J., Harvey, M. S., Kottelat, M., Kluge, N., Krell, F.-T., Kojima, J.-I., Kullander, S. O., Lucinda, P., Lyal, C. H. C., Pyle, R. L., Rheindt, F. E., Luisa Scioscia, C., Welter-Schultes, F., Whitmore, D., Yanega, D., Zhang, Z.-Q., Zhou, H.-Z. & Pape, T. (2023) Renaming taxa on ethical grounds threatens nomenclatural stability and scientific communication: Communication from the International Commission on Zoological Nomenclature. Zoological Journal of the Linnean Society, 127, 283–286.

Harvey, M. S. (2023a) A new species of Synsphyronus (Pseudoscorpiones: Garypidae) from eastern Australia. Australian Journal of Taxonomy, 18, 1–5.

Harvey, M. S. (2023b) A preliminary phylogeny for the pseudoscorpion family Garypinidae (Pseudoscorpiones: Garypinoidea), with new taxa and remarks on the Australasian fauna. Invertebrate Systematics, 37, 623–676.

Harvey, M. S., Burger, M. A. A., Abrams, K. M., Finston, T. L., Huey, J. A. & Perina, G. (2023a) The systematics of the pseudoscorpion genus Indohya (Pseudoscorpiones: Hyidae) in Australia. Zootaxa, 5342, 1–119.

Harvey, M. S., Wilson, J. D. & Rix, M. G. (2023b) Three new species of open-holed trapdoor spider genus Kwonkan (Mygalomorphae: Anamidae) from central Australia. Australian Journal of Taxonomy, 21, 1–9.

Harvey, M. S., Wilson, J. D. & Rix, M. G. (2023c) Two new species of the open-holed trapdoor spider genus Proshermacha (Araneae: Mygalomorphae: Anamidae) from southern Western Australia. Australian Journal of Taxonomy, 43, 1–13.

Hlebec, D. & Harvey, M. S. (2023) Lažištipavci – zašto tako malo znamo? Subterranea Croatica, 21, 47–53.

Leenders, X. J. R., Beach, D. & Harvey, M. S. (2023) A new species of open-holed trapdoor spider of the genus Proshermacha (Mygalomorphae: Anamidae) from Western Australia. Australian Journal of Taxonomy, 20, 1–10.

Marsh, J. R., Milner, S. J., Shaw, M., Stempel, A., Harvey, M. S. & Rix, M. G. (2023) A case for below ground dispersal? Insights into the biology, ecology and conservation of blind cave spiders in the genus Troglodiplura (Mygalomorphae: Anamidae). Insects, 14, 1–22.

Novak, J., Harvey, M. S., Szabó, M., Hammel, J., Harms, D., Kotthoff, U., Hörweg, C., Brazidec, M. & Ősi, A. (2023) A new Mesozoic record of the pseudoscorpion family Garypinidae from Upper Cretaceous (Santonian) Ajkaite amber, Ajka area, Hungary. Cretaceous Research, 153, 1–9.

Rheindt, F. E., Bouchard, P., Pyle, R. L., Welter-Schultes, F., Aescht, E., Ahyong, S. T., Ballerio, A., Bourgoin, T., Ceríaco, L. M. P., Dmitriev, D., Evenhuis, N., Grygier, M. J., Harvey, M. S., Kottelat, M., Kluge, N., Krell, F.-T., Kojima, J.-i., Kullander, S. O., Lucinda, P., Lyal, C. H. C., Scioscia, C. L., Whitmore, D., Yanega, D., Zhang, Z.-Q., Zhou, H.-Z. & Pape, T. (2023) Tightening the requirements for species diagnoses would help integrate DNA-based descriptions in taxonomic practice. PloS Biology, 21, e3002251.

Rix, M. G., Wilson, J. D. & Harvey, M. S. (2023a) A revision of the open-holed trapdoor spiders of the Teyl damsonoides-group (Mygalomorphae: Anamidae: Teylinae) from south-western Australia. Journal of Arachnology, 53, 287–377.

Rix, M. G., Wilson, J. D., Laidlaw, M. J., Harvey, M. S., Rix, A. G. & Rix, D. C. (2023b) Demography, passive surveillance and potential habitat modelling of an Australian giant trapdoor spider (Idiopidae: Euoplos grandis) from the Queensland Brigalow Belt: half a decade of population monitoring for conservation outcomes. Austral Entomology, 62, 200–219.

Rogers, D. J., Alderman, A., Barrett, S., Buchan, A., Burbidge, A. H., Churchill, T., Comer, S., Gales, R., Geary, W., Gosper, C. R., Harvey, M. S., Higham, J., Leonard, S., Lloyd, H., Loan, L., Page, M., Reid, T. & Walker, I. S. (2023) State agency responses to support biodiversity recovery follwoing the 2019–2020 wildfires. In: L. Rumpff, S. Legge, S. van Leeuwen, B. Wintle & J. Woinarski (Eds), Australia's megafires: biodiversity impacts and lessons from 2019–2020. CSIRO Publishing, Melbourne, pp. 388–404.

Romero-Ortiz, C., Samiento, C. E. & Harvey, M. S. (2023) A new genus and five new species of false scorpions (Arachnida, Pseudoscorpiones, Withiidae) from Colombia. ZooKeys, 1184, 301–326.

Stanczak, N., Harvey, M. S., Harms, D., Hammel, J. U., Kotthoff, U. & Loria, S. F. (2023) A new pseudoscorpion genus (Garypinoidea: Garypinidae) from the Eocene supports extinction and range contraction in the European paleobiota. PeerJ, 11, 1–27.

Wilson, J. D., Bond, J. E., Harvey, M. S., Ramiréz, M. G. & Rix, M. G. (2023a) Correlation with a limited set of behavioral niches explains the convergence of somatic morphology in mygalomorph spiders. Ecology and Evolution, 13, 1-17.

Wilson, J. D., Rix, M. G. & Harvey, M. S. (2023b) Description of five new Aname L. Koch, 1873 (Araneae, Anamidae) species collected on Bush Blitz expeditions. European Journal of Taxonomy, 890, 1–22.

Wilson, J. D., Rix, M. G. & Harvey, M. S. (2023c) A new spider species of the genus Proshermacha (Mygalomorphae: Anamidae) from the Coolgardie and Murchison bioregions of Western Australia, collected on a Bush Blitz expedition. Australian Journal of Taxonomy, 19, 1–7.

2022

Amieva Mau, S., Harvey, M. S. & Harms, D. (2022) New syarinid pseudoscorpions from Ecuador (Pseudoscorpiones, Syarinidae). European Journal of Taxonomy, 821, 102–149.

Ballesteros, J. A., Santibáñez-López, C. E., Baker, C. M., Benavides Silva, L. R., Cunha, T. J., Gainett, G., Ontano, A. Z., Setton, E. V. W., Arango, C. P., Gavish-Regev, E., Harvey, M. S., Wheeler, W. C., Hormiga, G., Giribet, G. & Sharma, P. P. (2022) Comprehensive species sampling and sophisticated algorithmic approaches refute the monophyly of Arachnida. Molecular Biology and Evolution, 39, 1–16.

Geißler, C., Kotthoff, U., Hammel, J., Harvey, M. S. & Harms, D. (2022) The first fossil of the pseudoscorpion family Ideoroncidae (Arachnida: Pseudoscorpiones): a new taxon from the mid-Cretaceous of norarabithern Myanmar. Cretaceous Research, 130, 1–12.

Harms, D., Nentwig, W., Gloor, D. & Harvey, M. S. (2022) Introducing the World Arachnida Catalog: the new research environment for (almost all) arachnid orders. Journal of Arachnology, 50, 113–117.

Harvey, M. S. (2022) Three new species of the pseudoscorpion genus Synsphyronus (Pseudoscorpiones: Garypidae) from semi-arid Western Australia. Australian Journal of Taxonomy, 5, 1–15.

Harvey, M. S., Bouchard, P., Krell, F.-T. & Rheindt, F. E. (2022a) Election of new ICZN Commissioners using an on-line platform. Bulletin of Zoological Nomenclature, 79, 6–7.

Harvey, M. S. & Harms, D. (2022) The pseudoscorpion genus Centrochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) from central Asia and description of a new species from Nepal. Journal of Arachnology, 50, 158–174.

Harvey, M. S., Wilson, J. D. & Rix, M. G. (2022b) Two new species of the open-holed trapdoor spider genus Aname (Araneae: Mygalomorphae: Anamidae) from southern Australia. Australian Journal of Taxonomy, 2, 1–8.

Lorenz, M., Loria, S. F., Harvey, M. S. & Harms, D. (2022) The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the “hottest” biodiversity hotspots. Arthropod Systematics & Phylogeny, 80, 649–691.

Marimon, K., Villareal B., E. & Harvey, M. S. (2022) Two new species of Rhopalochernes Beier, 1932 (Pseudoscorpiones: Chernetidae) from Colombia. Zootaxa, 5150, 397–410.

Pepato, A. R., Costa, S. G. d. S., Harvey, M. S. & Klimov, P. B. (2022) One-way ticket to the blue: A large-scale, dated phylogeny revealed asymmetric land-to-water transitions in acariform mites (Acari: Acariformes). Molecular Phylogenetics and Evolution, 107626, 1–15.

Rix, M. G. & Harvey, M. S. (2022) A new species of the spiny trapdoor spider genus Eucyrtops (Mygalomorphae: Idiopidae) from south-western Australia. Australian Journal of Taxonomy, 4, 1–6.

Rix, M. G., Worthington Wilmer, J. & Harvey, M. S. (2022) Rainforest pelican spiders (Archaeidae: Austrarchaea) of south-eastern Queensland, Australia: two new species and a distributional reassessment of regional endemic clades. Journal of Arachnology, 50, 231–249.

Santibáñez-López, C. E., Aharon, S., Ballesteros, J. A., Gainett, G., Baker, C. M., González-Santillán, E., Harvey, M. S., Hassan, M. K., Abu Almaaty, A. H., Aldeyarbi, S. M., Monod, L., Ojanguren-Affilastro, A., Pinto-da-Rocha, R., Zvik, Y., Gavish-Regev, E. & Sharma, P. P. (2022) Phylogenomics of scorpions reveal contemporaneous diversification of scorpion mammalian predators and mammal-active sodium channel toxins. Systematic Biology, 71, 1281–1289.

Thiele, K. R., Harvey, M. S., Hutchings, P., May, T. W., Melville, J. & Travouillon, K. J. (2022) Introducing the Australian Journal of Taxonomy, a new, fully-online, fully open-access journal for the rapid publication of new Australian species and other taxa. Australian Journal of Taxonomy, 1, 1–7.

Wilson, J. D., Harvey, M. S. & Rix, M. G. (2022) Euoplos eungellaensis, sp. nov. (Idiopidae), a new golden trapdoor spider from central-eastern Queensland. Australian Journal of Taxonomy, 5, 1–8.

You, J. Y., Yoo, J.-S., Harvey, M. S. & Harms, D. (2022) Some cryptic Korean karst creatures: Revalidation of the pseudoscorpion genus Spelaeochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) and description of two new species from Korea. Journal of Arachnology, 50, 135–157.

2021

Cullen, K. L. & Harvey, M. S. (2021a) New species of the pseudoscorpion genus Synsphyronus (Pseudoscorpiones: Garypidae) from Australia. Records of the Western Australian Museum, 36, 33–65.

Cullen, K. L. & Harvey, M. S. (2021b) Two new species of the pseudoscorpion genus Geogarypus (Pseudoscorpiones: Geogarypidae) from northern Australia. Records of the Western Australian Museum, 36, 71–78.

Harvey, M. S. (2021a) A new genus of the pseudoscorpion family Chernetidae (Pseudoscorpiones) from southern Australia with Gondwanan affinities. Journal of Arachnology, 48, 300–310.

Harvey, M. S. (2021b) A new species of Garypus (Pseudoscorpiones: Garypidae) from southern Thailand. Revue Suisse de Zoologie, 128, 221–225.

Harvey, M. S. & Cullen, K. L. (2021) A new species of the pseudoscorpion genus Anatemnus (Pseudoscorpiones: Atemnidae) from tropical Australia. Records of the Western Australian Museum, 36, 66–70.

Marek, P., Buzatto, B., Shear, W. A., Means, J. C., Black, D. G., Harvey, M. S. & Rodriguez, J. (2021) The first true millipede—1,306 legs long. Scientific Reports, 11, 1–8.

Ontano, A. Z., Gainett, G., Aharon, S., Ballesteros, J. A., Benavides, L. R., Corbett, K. F., Gavish-Regev, E., Harvey, M. S., Monsma, S., Santibáñez-López, C. E., Setton, E. V. W., Zehms, J. T., Zeh, J. A., Zeh, D. W. & Sharma, P. P. (2021) Taxonomic sampling and rare genomic changes overcome long-branch attraction in the phylogenetic placement of pseudoscorpions. Molecular Biology and Evolution, 38, 2446–2467.

Rix, M. G., Wilson, J. D., Huey, J. A., Hillyer, M. J., Gruber, K. & Harvey, M. S. (2021a) Diversification of the mygalomorph spider genus Aname (Araneae: Anamidae) across the Australian arid zone: tracing the evolution and biogeography of a continent-wide radiation. Molecular Phylogenetics and Evolution, 160, 1–16.

Rix, M. G., Wood, H. M., Harvey, M. S. & Michalik, P. (2021b) Micro-computed tomography reveals a remarkable twin intromittent organ in spiders – a novelty for arachnids with direct sperm transfer. Frontiers in Ecology And Evolution, 9, 1–7.

Ward, M., Carwardine, J., Yong, C. J., Watson, J. E. M., Silcock, J., Taylor, G. S., Lintermans, M., Gillespie, G. R., Garnett, S. T., Woinarski, J., Tingley, R., Fensham, R. J., Hoskin, C. J., Hines, H. B., Roberts, J. D., Kennard, M. J., Harvey, M. S., Chapple, D. G. & Reside, A. E. (2021) A national-scale dataset for threats impacting Australia’s imperiled flora and fauna. Ecology and Evolution, 11, 11749–11761.

Wreidt, A.-L., Harvey, M. S., Hammel, J. U., Kotthoff, U. & Harms, D. (2021) The second chthonioid pseudoscorpion (Pseudoscorpiones: Chthoniidae) from mid-Cretaceous Burmese amber: a new genus with unique morphological features and potential Gondwanan affinities. Journal of Arachnology, 48, 311–321.

2020

Harvey, M.S., Gruber, K., Hillyer, M.J. and Huey, J.A. (2020). Five new species of the open-holed trapdoor spider genus Aname (Araneae: Mygalomorphae: Anamidae) from Western Australia, with a revised generic placement for Aname armigera. Records of the Western Australian Museum 35: 10–38.

Harvey, M.S., Rix, M.G., Hillyer, M.J. and Huey, J.A. (2020). The systematics and phylogenetic position of the troglobitic Australian spider genus Troglodiplura (Araneae: Mygalomorphae), with a new classification for Anamidae. Invertebrate Systematics 34: 799–822.

Harvey, M.S. and Evenhuis, N.L. (2020). Revised authorship and dates of some genus-group names in Arachnida proposed without type species after 1930. Arachnology 18: 623–628.

Harvey, M.S. and Cullen, K.L. (2020). A remarkable new troglobitic species of Parobisium (Pseudoscorpiones: Neobisiidae) from California. Arachnology 18: 591–596.

Abrams, K.M., Huey, J.A., Hillyer, M.J., Didham, R.K. and Harvey, M.S. (2020). A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara. Zootaxa 4864: 1–75.

Rix, M.G., Wilson, J.D. and Harvey, M.S. (2020). The open-holed trapdoor spiders (Mygalomorphae: Anamidae: Namea) of Australia’s D’Aguilar Range: revealing an unexpected subtropical hotspot of rainforest diversity. Zootaxa 4861: 71–91.

Castalanelli, M.A., Framenau, V.W., Huey, J.A. and Harvey, M.S. (2020). New species of the open-holed trapdoor spider genus Aname (Araneae: Mygalomorphae: Anamidae) from arid Western Australia. Journal of Arachnology 48: 169–213.

Rix, M.G., Wilson, J.D. and Harvey, M.S. (2020). First phylogenetic assessment and taxonomic synopsis of the open-holed trapdoor spider genus Namea (Mygalomorphae: Anamidae): a highly diverse mygalomorph lineage from Australia’s tropical eastern rainforests. Invertebrate Systematics 34: 679–726.

Harvey, M.S. (2020). Synsphyronus platnicki sp. nov.: first Synsphyronus (Pseudoscorpiones: Garypidae) from New Caledonia. Arachnology 18: 468–472.

Harvey, M.S., Hillyer, M.A., Carvajal, J.I. and Huey, J.A. (2020). Supralittoral pseudoscorpions of the genus Garypus (Pseudoscorpiones: Garypidae) from the Indo-West Pacific region, with a review of the subfamily classification of Garypidae. Invertebrate Systematics 34: 34—87.

2019

Abrams, K.M., Huey, J.A., Hillyer, M.A., Humphreys, W.F., Didham, R.K. and Harvey, M.S. (2019). Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution. 139 (106532): 1–12.

Benavides, L.R., Cosgrove, J.G., Harvey, M.S. and Giribet, G. (2019). Phylogenomic interrogation resolves the backbone of the Pseudoscorpiones Tree of Life. Molecular Phylogenetics and Evolution. 139 (106509): 1–14.

Huey, J.A., Rix, M.G., Wilson, J.D., Hillyer, M.J. and Harvey, M.S. (2019). Open-holed trapdoor spiders of the genus Teyl (Mygalomorphae: Nemesiidae: Anamini) from Western Australia’s Pilbara bioregion: a new species and expanded phylogenetic assessment. Zootaxa 4674: 349–362.

Harms, D., Roberts, J.D. and Harvey, M.S. (2019). Climate variability impacts on diversification processes in a biodiversity hotspot: a phylogeography of ancient pseudoscorpions in southwestern Australia. Zoological Journal of the Linnean Society 186: 934–949.

Huey, J.A., Hillyer, M.J. and Harvey, M.S. (2019). Phylogenetic relationships and biogeographic history of the Australian trapdoor spider genus Conothele (Araneae: Mygalomorphae: Halonoproctidae): diversification into arid habitats in an otherwise tropical radiation. Invertebrate Systematics 33: 628–643.

Ott, R., Ubick, D., Bonaldo, A.B., Brescovit, A.D. and Harvey, M.S. (2019). A revision of the New World goblin spider genus Cinetomorpha Simon, 1892 revalidated from Gamasomorpha Karsch, 1881 (Araneae, Oonopidae, Oonopinae). Zootaxa 4641: 1–152.

Car, C.A., Harvey, M.S. and Huey, J.A. (2019). The millipede genus Antichiropus (Diplopoda: Polydesmida: Paradoxosomatidae), part 3: species of the Pilbara bioregion of Western Australia. Zootaxa 4617: 1–71.

Gibson, L., Humphreys, W.F., Harvey, M.S., Hyder, B. and Winzer, A. (2019). Shedding light on the hidden world of subterranean fauna: a transdisciplinary research approach. Science of the Total Environment 684: 381–389.

Rix, M.G., Wilson, J.D., Rix, A.G., Wojcieszek, A.M., Huey, J.A. and Harvey, M.S. (2019). Population demography and biology of a giant spiny trapdoor spider (Idiopidae: Euoplos) from inland Queensland: developing a ‘slow science’ study system to address a conservation crisis. Austral Entomology 58: 282–297.

Novák, J. and Harvey, M.S. (2019). The status of the pseudoscorpion genus Indogarypus Beier, 1957 with new descriptions of three species of Geogarypus Chamberlin, 1930 (Arachnida: Pseudoscorpiones: Geogarypidae). Journal of Arachnology 47: 124–131.

Johnson, J., Romero-Ortiz, C., Mathew, A.V., Sebastian, P.A., Joseph, M.M. and Harvey, M.S. (2019). A review of the pseudoscorpion genus Metawithius (Pseudoscorpiones: Withiidae) from the Indian subcontinent. Journal of Arachnology 47: 84–94.

Rix, M.G., Wilson, J.D. and Harvey, M.S. (2019). A revision of the white-headed spiny trapdoor spiders of the genus Euoplos (Mygalomorphae: Idiopidae: Arbanitinae): a remarkable lineage of rare mygalomorph spiders from the south-western Australian biodiversity hotspot. Journal of Arachnology 47: 63–76.

Harvey, M.S. and Mahnert, V. (2019). The status of the Central American pseudoscorpion genus Coprochernes Beier, a new synonym of Neoallochernes Hoff (Pseudoscorpiones: Chernetidae). Arachnology 18: 7–13.

Romero-Ortiz, C. and Harvey, M.S. (2019). The pseudoscorpion genus Verrucachernes (Pseudoscorpiones: Chernetidae) in the Indian region. Zootaxa 4569: 337–346.

Kropf, C., Blick, T., Brescovit, A.D., Chatzaki, M., Dupérré, N., Gloor, D., Haddad, C.R., Harvey, M.S., Jäger, P., Marusik, Y.M., Ono, H., Rheims, C.A. and Nentwig, W. (2019). How not to delimit taxa: a critique on a recently proposed “pragmatic classification” of jumping spiders (Arthropoda: Arachnida: Araneae: Salticidae). Zootaxa 4545: 444–446.